Frank Ruddle

Frank Ruddle
Frank Ruddle between Ruth Kirschstein and Donald S. Fredrickson in 1978
Born
Francis Hugh Ruddle

(1929-08-09)August 9, 1929
DiedMarch 10, 2013(2013-03-10) (aged 83)
Alma materUniversity of California, Berkeley (PhD)
AwardsDickson Prize in Medicine (1982)
William Allan Award (1984)
Scientific career
InstitutionsYale University
ThesisChromosome variation in cell populations (1960)
Doctoral studentsAnne Ferguson-Smith

Francis Hugh Ruddle (1929–2013) was an American cell and developmental biologist who was the Sterling Professor at Yale University. Ruddle was an early visionary of the Human Genome Project and created the first genetically modified mouse. He was a pioneer in both human and mouse genetics.