Pseudosuchia

Pseudosuchia, from Ancient Greek ψεύδος (pseúdos), meaning "false", and σούχος (soúkhos), meaning "crocodile" is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs", in contrast to the "bird-line archosaurs" or Avemetatarsalia. Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.

The clade Pseudosuchia is potentially equivalent to another term, Crurotarsi, even though the latter has a different, node-based definition: "all taxa the least inclusive clade containing Rutiodon carolinensis (Emmons, 1856), and Crocodylus niloticus (Laurenti, 1768)." Many paleontologists of the late 20th century took this proposal for granted, using Crurotarsi as the term for crocodilian ancestors. In 2011, a major revision of Triassic archosaur relations proposed that Rutiodon's group, Phytosauria, was not as closely related to other traditional "crurotarsans", at least compared to avemetatarsalians such as pterosaurs and dinosaurs. Under that interpretation, Crurotarsi would be a much broader clade than Pseudosuchia. Other recent studies continue to support a more traditional phylogeny with phytosaurs as an early branch of Pseudosuchia. If the traditional interpretation is maintained, Pseudosuchia and Crurotarsi are roughly equivalent categories.

Contrary to popular belief, crocodilians differ significantly from their ancestors and distant relatives, as Pseudosuchia contains a staggering diversity of reptiles with many different lifestyles. Early pseudosuchians were successful in the Triassic period. They included giant, quadrupedal apex predators such as Saurosuchus, Prestosuchus, and Fasolasuchus. Ornithosuchids were large scavengers, while erpetosuchids and gracilisuchids were small, light-footed predators. A few groups acquired herbivorous diets, such as the heavily armored aetosaurs, and several were bipedal, such as Poposaurus and Postosuchus. The bizarre, ornithomimid-like shuvosaurids were both bipedal and herbivorous, with toothless beaks.

Many of these Triassic pseudosuchian groups went extinct at or before the Triassic–Jurassic extinction event. However, one group, the crocodylomorphs, survived the major extinction. Crocodylomorphs themselves evolved a diverse array of lifestyles during the Jurassic and Cretaceous periods, although living pseudosuchians only include crocodiles, alligators, caimans, and gavialids.