Tyrannosaurus

Tyrannosaurus
Temporal range: Late Cretaceous (Latest Campanian?-Maastrichtian), Possible Middle Campanian record
Reconstruction of the T. rex type specimen at the Carnegie Museum of Natural History
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Superfamily: Tyrannosauroidea
Family: Tyrannosauridae
Tribe: Tyrannosaurini
Genus: Tyrannosaurus
Osborn, 1905
Type species
Tyrannosaurus rex
Osborn, 1905
Other species
Synonyms
Genus synonymy
  • Dinotyrannus
    Olshevsky, 1995
  • Dynamosaurus
    Osborn, 1905
  • Manospondylus
    Cope, 1892
  • Nanotyrannus?
    Bakker, Williams & Currie, 1988
  • Stygivenator
    Olshevsky, 1995
Species synonymy
  • Aublysodon amplus?
    Marsh, 1892
  • Deinodon amplus?
    (Marsh, 1892) Hay, 1902
  • Manospondylus amplus?
    (Marsh, 1892) Olshevsky, 1978
  • Stygivenator amplus?
    (Marsh, 1892) Olshevsky, 1995
  • Tyrannosaurus amplus?
    (Marsh, 1892) Hay, 1930
  • Aublysodon cristatus?
    Marsh, 1892
  • Deinodon cristatus?
    (Marsh, 1892) Hay, 1902
  • Stygivenator cristatus?
    (Marsh, 1892) Olshevsky, 1995
  • Ornithomimus grandis
    Marsh, 1890
  • Manospondylus gigas
    Cope, 1892
  • Dynamosaurus imperiosus
    Osborn, 1905
  • Tyrannosaurus imperiosus
    (Osborn, 1905) Swinton, 1970
  • Gorgosaurus lancensis
    Gilmore, 1946
  • Albertosaurus lancensis
    (Gilmore, 1946) Russell, 1970
  • Deinodon lancensis
    (Gilmore, 1946) Kuhn, 1965
  • Aublysodon lancensis
    (Gilmore, 1946) Charig in Appleby, Charig, Cox, Kermack & Tarlo, 1967
  • Nanotyrannus lancensis
    (Gilmore, 1946) Bakker, Williams & Currie, 1988
  • Albertosaurus "megagracilis"
    Paul, 1988a (nomen nudum)
  • Dinotyrannus megagracilis
    Olshevsky, 1995
  • Aublysodon molnaris
    Paul, 1988a
  • Aublysodon molnari
    Paul, 1988a emend Paul, 1990
  • Stygivenator molnari
    (Paul, 1988a emend Paul, 1990) Olshevsky, 1995

Tyrannosaurus (/tɪˌrænəˈsɔːrəs, t-/) is a genus of large theropod dinosaur. The type species Tyrannosaurus rex (rex meaning 'king' in Latin), often shortened to T. rex or colloquially t-rex, is one of the best represented theropods. It lived throughout what is now western North America, on what was then an island continent known as Laramidia. Tyrannosaurus had a much wider range than other tyrannosaurids. Fossils are found in a variety of geological formations dating to the latest Campanian-Maastrichtian ages of the late Cretaceous period, 72.7 to 66 million years ago, with isolated specimens possibly indicating an earlier origin in the middle Campanian. It was the last known member of the tyrannosaurids and among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event.

Like other tyrannosaurids, Tyrannosaurus was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the forelimbs of Tyrannosaurus were short but unusually powerful for their size, and they had two clawed digits. The most complete specimen measures 12.3–12.4 m (40–41 ft) in length, but according to most modern estimates, Tyrannosaurus could have exceeded sizes of 13 m (43 ft) in length, 3.7–4 m (12–13 ft) in hip height, and 8.8 t (8.7 long tons; 9.7 short tons) in mass. Although some other theropods might have rivaled or exceeded Tyrannosaurus in size, it is still among the largest known land predators, with its estimated bite force being the largest among all terrestrial animals. By far the largest carnivore in its environment, Tyrannosaurus rex was most likely an apex predator, preying upon hadrosaurs, juvenile armored herbivores like ceratopsians and ankylosaurs, and possibly sauropods. Some experts have suggested the dinosaur was primarily a scavenger. The question of whether Tyrannosaurus was an apex predator or a pure scavenger was among the longest debates in paleontology. Most paleontologists today accept that Tyrannosaurus was both a predator and a scavenger.

Some specimens of Tyrannosaurus rex are nearly complete skeletons. Soft tissue and proteins have been reported in at least one of these specimens. The abundance of fossil material has allowed significant research into many aspects of the animal's biology, including its life history and biomechanics. The feeding habits, physiology, and potential speed of Tyrannosaurus rex are a few subjects of debate. Its taxonomy is also controversial. The Asian Tarbosaurus bataar is very closely related to Tyrannosaurus and has sometimes been seen as a species of this genus. Several North American tyrannosaurids have been synonymized with Tyrannosaurus, while some Tyrannosaurus specimens have been proposed as distinct species. The validity of these species, such as the more recently discovered T. mcraeensis, is contentious.

As the archetypal theropod, Tyrannosaurus has been one of the best-known dinosaurs since the early 20th century and has been featured in film, advertising, postal stamps, and many other media.