Wing coupling

Some four-winged insect orders, such as the Lepidoptera, have developed a wide variety of morphological wing coupling mechanisms in the imago which render these taxa as "functionally dipterous" (effectively two-winged) for efficient insect flight. All but the most basal forms exhibit this wing coupling.: 4266 

The mechanisms are of three different types - jugal, frenulo-retinacular and amplexiform.

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