Massospondylus

Massospondylus (/ˌmæsoʊˈspɒndɪləs/ MASS-oh-spon-di-lus) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Although the original fossils were destroyed in London during a bombing raid in World War II, a plethora of specimens have since been assigned to the genus, making it one of the best-known sauropodomorphs from the Early Jurassic. The genus lived from the Hettangian to Pliensbachian ages, ca. 200–183 million years ago. Most fossils come from the upper Elliot and Clarens formations of South Africa and Lesotho, but the genus is also found in the Forest Sandstone and the Mpandi Formation of Zimbabwe. Material from the US, India, and Argentina was previously assigned to the genus, but the US and Argentinian specimens are now assigned to their own genera (Sarahsaurus and Adeopapposaurus). Because of their great abundance, Massospondylus fossils have been used to date rocks, and a biozone, the Massospondylus Assemblage Zone, is named after it.

The type species is M. carinatus; seven other species have been named during the past 150 years, but among these, only M. kaalae is still considered valid. Originally, Massospondylus and similar dinosaurs have been regarded as theropods, but are now classified as basal ("early diverging") members of Sauropodomorpha. This group also includes sauropods. Within sauropodomorphs, Massospondylus is often classified in the family Massospondylidae. The genus was 4–6 metres (13–20 ft) long, and had a long neck and tail, a small head, and a slender body. It is distinguished from related genera by the very elongated vertebrae of the front portion of the neck, amongst other features. Although Massospondylus was long depicted as quadrupedal (four-legged), it is now considered to have been bipedal (two-legged).

It was probably a plant eater (herbivore), although some have speculated that early sauropodomorphs may have been omnivorous. On each of its hands, it bore a sharp thumb claw that was probably used in feeding, possibly to uproot vegetation or to pull down branches while rearing up. Clutches with eggs have been found, some of which contained embryos; these are among the oldest eggs and embryos of an amniote in the fossil record. The eggshell was extremely thin (ca. 0.1 mm), unlike the much thicker eggshells in later dinosaurs. The embryos had longer arms than adults and a proportionally very large head, leading researchers to suggest that they were quadrupedal and shifted to a bipedal posture later during growth. Newer research instead suggested that Massospondylus was bipedal at all ages. Individuals accelerated or slowed down their growth depending on environmental factors such as food availability. The oldest known specimen was around 20 years of age.