Pachycephalosaurus

Pachycephalosaurus
Temporal range: Late Cretaceous (Maastrichtian),
Cast of the "Sandy" specimen, Royal Ontario Museum
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Pachycephalosauria
Family: Pachycephalosauridae
Tribe: Pachycephalosaurini
Genus: Pachycephalosaurus
Brown & Schlaikjer, 1943
Type species
Pachycephalosaurus grangeri
Brown & Schlaikjer, 1943
(Junior synonym of P. wyomingensis)
Species
  • P. wyomingensis
    (Gilmore, 1931) (conserved name)
  • P. spinifer?
    (Galton & Sues, 1983)
Synonyms
Synonyms of P. wyomingensis
  • Tylosteus ornatus
    Leidy, 1872 (rejected name)
  • Troodon wyomingensis
    Gilmore, 1931
  • Pachycephalosaurus grangeri
    Brown & Schlaikjer, 1943
  • Pachycephalosaurus reinheimeri
    Brown & Schlaikjer, 1943
  • Stenotholus kohleri
    Giffin, Gabriel & Johnson, 1988
  • Stygimoloch spinifer?
    Galton & Sues, 1983
  • Dracorex hogwartsia?
    Bakker et al., 2006

Pachycephalosaurus (/ˌpækɪˌsɛfələˈsɔːrəs/; meaning "thick-headed lizard", from Greek pachys-/παχύς- "thickness", kephalon/κεφαλή "head" and sauros/σαῦρος "lizard") is a genus of pachycephalosaurid ornithischian dinosaur. The type species, P. wyomingensis, is the only known definitive species. The possibly synonymous taxon, Stygimoloch, might represent a distinct genus or a second species, P. spinifer. It lived during the Maastrichtian age of the Late Cretaceous period in what is now western North America. Remains have been excavated in Montana, South Dakota, Wyoming, and Alberta. Mainly known from a single skull and a few extremely thick skull roofs (at 22 cm or 9 in thick), Pachycephalosaurus is estimated to have reached 4.5 m (15 ft) long and weighed 370–450 kg (820–990 lb). More complete fossils would come to be found in the following years.

Pachycephalosaurus was among the last species of non-avian dinosaurs on Earth before the Cretaceous–Paleogene extinction event. The genus Tylosteus has been synonymized with Pachycephalosaurus, as have the genera Stygimoloch and Dracorex, in recent studies.

Like other pachycephalosaurids, Pachycephalosaurus was a bipedal herbivore, possessing long, strong legs and somewhat small arms with five-fingered hands. Pachycephalosaurus is the largest-known pachycephalosaur, known for having an extremely thick, slightly domed skull roof; visually, the structure of the skull suggests a "battering ram" function in life, evolved for use as a defensive mechanism or intra-species combat, similar to what is seen with today's bighorn sheep or muskoxen (with male animals routinely charging and head-butting each other for dominance). This hypothesis has actually been highly disputed in recent years.