Tusk shell

Scaphopods
Temporal range: CarboniferousNeogene (
Various Scaphopoda, from left to right: Fissidentalium, Gadilida, Gadila, and Gadilida.
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Subphylum: Conchifera
Class: Scaphopoda
Bronn, 1862
Orders

Scaphopoda /skæˈfɒpədə/ (plural scaphopods /ˈskæfəpɒdz/, from Ancient Greek σκᾰ́φης skáphē "boat" and πούς poús "foot"), whose members are also known as tusk shells or tooth shells, are a class of shelled marine invertebrates belonging to the phylum Mollusca with worldwide distribution and are the only class of exclusively infaunal marine molluscs. Shells of species within this class range in length 0.5–18 cm (0.20–7.09 in) (with Fissidentalium metivieri as the longest). Members of the order Dentaliida tend to be larger than those of the order Gadilida.

These molluscs live in soft substrates offshore (usually not intertidally). Because of this subtidal habitat and the small size of most species, many beachcombers are unfamiliar with them; their shells are not as common or as easily visible in the beach drift as the shells of sea snails and clams.

Molecular data suggest that the scaphopods are a sister group to the cephalopods, although higher-level molluscan phylogeny remains unresolved.